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"Repetition sometimes works in poetry, but rarely in prose. The musical provocateur John Cage once wrote a lecture in which a single page was repeated fourteen times, with the refrain "If anybody is sleep let him go to sleep" (Cage, 1961). Midway through, the artist Jean Reynal stood up and screamed, "John, I dearly love you, but I can't bear another minute.
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In essence, the bee's azimuth system acts like a sundial run in reverse, and like a sundial, it has to be calibrated. A sundial, which must be oriented with respect to a known compass direction, calculates the time of day based on where the sun is; the navigational centers in the bee's brain calculate where the sun should be based on the time of day. As a consequence, the one thing that bees can't cope with is the discalibration that results from jet lag. In a famous 1960s experiment, Max Renner packed up a hive of bees in Long Island, New York, flew them to Davis, California, and tested their ability to navigate with the sun as a landmark. The jetlagged bees consistently misoriented themselves by 45 degrees, precisely as though they believed it was three hours later. The complex circuitry that allows the bee to use the sun as a guide is built in, but it is not that genes trump the environment (or the other way around). Instead, genes enable creatures to make sensible use of their particular environment. Learning is not the antithesis of innateness but one of its most important products.
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If there is not preformation, and no blueprint, there is also no getting away from the environment. Genes do not guarantee particular products; rather, they provide particular options: To every gene there is an IF, and with that IF comes an option. In many cases, those options are selected based on cues from the environment, and it is for that reason, more than any other, that the answer to the nature-nurture question is not one or the other, but both.