The student of development finds, not only that the chick commences its existence as an egg, primarily identical, in all essential respects, with that of the Dog, but that the yelk of this egg undergoes division—that the primitive groove arises, and that the contiguous parts of the germ are fashioned, by precisely similar methods into a young chick, which, at one stage of its existence, is so like the nascent Dog, that ordinary inspection would hardly distinguish the two. The history of the development of any other vertebrate animal, Lizard, Snake, Frog, or Fish, tells the same story.

Within the last half century, the labours of such men as Von Baer, Rathke, Reichert, Bischof, and Remak, have almost completely unravelled... the successive stages of development which... are now as well known to the embryologist as are the steps of the metamorphosis of the silk-worm moth to the school boy.

I now propose briefly to... set forth, in a form intelligible to those who possess no special acquaintance with anatomical science, the chief facts upon which all conclusions respecting the nature and the extent of the bonds which connect man with the brute world must be based: I shall then indicate the one immediate conclusion which, in my judgment, is justified by those facts, and I shall finally discuss the bearing of that conclusion upon the hypotheses which have been entertained respecting the Origin of Man.

In a well worn metaphor, a parallel is drawn between the life of man and the metamorphosis of the caterpillar into the butterfly; but the comparison may be more just as well as more novel, if for its former term we take the mental progress of the race. History shows that the human mind, fed by constant accessions of knowledge, periodically grows too large for its theoretical coverings, and bursts them asunder to appear in new habiliments, as the feeding and growing grub, at intervals, casts its too narrow skin and assumes another, itself but temporary. Truly the imago state of Man seems to be terribly distant, but every moult is a step gained, and of such there have been many.

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Each such answer to the great question, invariably asserted by the followers of its propounder, if not by himself, to be complete and final, remains in high authority and esteem, it may be for one century, or it may be for twenty: but, as invariably, Time proves each reply to have been a mere approximation to the truth—tolerable chiefly on account of the ignorance of those by whom it was accepted, and wholly intolerable when tested by the larger knowledge of their successors.

The sceptics end in the infidelity which asserts the problem to be insoluble, or in the atheism which denies the existence of any orderly progress and governance of things: the men of genius propound solutions which grow into systems of Theology or of Philosophy, or veiled in musical language which suggests more than it asserts, take the shape of the Poetry of an epoch.

Most of us, shrinking from the difficulties and dangers which beset the seeker after original answers to these riddles, are contented to ignore them altogether, or to smother the investigating spirit under the featherbed of respected and respectable tradition. But, in every age, one or two restless spirits, blessed with that constructive genius, which can only build on a secure foundation, or cursed with the mere spirit of scepticism, are unable to follow in the well-worn and comfortable track of their forefathers and contemporaries, and unmindful of thorns and stumbling-blocks, strike out into paths of their own.

Once in a generation, a Wallace may be found physically, mentally, and morally qualified to wander unscathed through the tropical wilds of America and of Asia; to form magnificent collections as he wanders; and withal to think out sagaciously the conclusions suggested by his collections: but, to the ordinary explorer or collector, the dense forests of equatorial Asia and Africa, which constitute the favourite habitation of the Orang, the Chimpanzee, and the Gorilla, present difficulties of no ordinary magnitude: and the man who risks his life by even a short visit to the malarious shores of those regions may well be excused if he shrinks from facing the dangers of the interior; if he contents himself with stimulating the industry of the better seasoned natives, and collecting and collating the more or less mythical reports and traditions with which they are too ready to supply him. In such a manner most of the earlier accounts of the habits of the man-like Apes originated...

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The whole analogy of natural operations furnishes so complete and crushing an argument against the intervention of any but what are termed secondary causes, in the production of all the phenomena of the universe; that, in view of the intimate relations between Man and the rest of the living world; and between the forces exerted by the latter and all other forces, I can see no excuse for doubting that all are co-ordinated terms of Nature's great progression, from the formless to the formed—from the inorganic to the organic—from blind force to conscious intellect and will.

The greater part of the substance of the following Essays has already been published in the form of Oral Discourses, addressed to widely different audiences, during the past three years. Upon the subject of the second Essay, I delivered six Lectures to the Working Men in 1860, and two, to the members of the Philosophical Institution of Edinburgh in 1862.