The live-and-let-live system that emerged in the bitter trench warfare of World War I demonstrates that friendship is hardly necessary for cooperatio… - Robert Axelrod

Once again, there is an important contrast between a zero-sum game like chess and a non-zero-sum game like the iterated Prisoner's Dilemma. In chess, it is useful to keep the other player guessing about your intentions. The more the other player is in doubt, the less efficient will be his or her strategy. Keeping one's intentions hidden is useful in a zero-sum setting where any inefficiency in the other player's behavior will be to your benefit. But in a non-zero-sum setting it does not always pay to be so clever. In the iterated Prisoner's Dilemma, you benefit from the other player's cooperation. The trick is to encourage that cooperation. A good way to do it is to make it clear that you will reciprocate. Words can help here, but as everyone knows, actions speak louder than words. That is why the easily understood actions of TIT FOR TAT are so effective.

In complex environments, individuals are not fully able to analyze the situation and calculate their optimal strategy. Instead they can be expected to adapt their strategy over time based upon what has been effective and what has not.

Thus cooperation can emerge even in a world of unconditional defection. The development cannot take place if it is tried only by scattered individuals who have no chance to interact with each other. But cooperation can emerge from small clusters of discriminating individuals, as long as these individuals have even a small proportion of their interactions with each other. Moreover, if nice strategies (those which are never the first to defect) come to be adopted by virtually everyone, then those individuals can afford to be generous in dealing with any others. By doing so well with each other, a population of nice rules can protect themselves against clusters of individuals using any other strategy just as well as they can protect themselves against single individuals. But for a nice strategy to be stable in the collective sense, it must be provocable. So mutual cooperation can emerge in a world of egoists without central control by starting with a cluster of individuals who rely on reciprocity.