Barriers of sea, or desert, or mountain, could never have been of the least avail, had the creative force acted independently of material laws, or had it not pleased the Author of Nature that the origin of new species should be governed by some secondary causes analogous to those which we see preside over the appearance of new varieties, which never appear except as the offspring of a parent stock very closely resembling them.

The only secondary cause... which has, as yet, been even conjecturally brought forward, to explain how, in the ordinary course of nature, a new specific form may be generated is, as Lamarck declared, 'variation,' and this has been rendered a far more probable hypothesis by the way in which Natural Selection is shown to give intensity and permanency to certain varieties.

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Hitherto no rival hypothesis has been proposed as a substitute for the doctrine of transmutation; for 'independent creation,' as it is often termed, or the direct intervention of the Supreme Cause, must simply be considered as an avowal that we deem the question to lie beyond the domain of science.

None of the observations are more in point, as bearing on the doctrine of what Hooker terms 'creation by variation,' than the great extent to which the internal characters and properties of plants, or their physiological constitution are capable of being modified, while they exhibit externally no visible departure from the normal form. ...When several of these internal or physiological modifications are accompanied by variation in size, habits of growth, colour of the flowers, and other external characters, and these are found to be constant in successive generations, botanists may well begin to differ in opinion as to whether they ought to regard them as distinct species or not.

Of Dr. Hooker, whom I have often cited in this chapter, Mr. Darwin has spoken in the Introduction to his 'Origin of Species, as one 'who had, for fifteen years, aided him in every possible way, by his large stores of knowledge, and his excellent judgement.' This distinguished botanist published his 'Introductory Essay to the Flora of Australia' in 1859, the year after the memoir on 'Natural Selection' was communicated to the Linnaean Society, and a few months before the appearance of the' Origin of Species.' ...no one was better qualified by observation and reflection to give an authoritative opinion on the question, whether the present vegetation of the globe is or is not in accordance with the theory which Mr. Darwin has proposed. We cannot but feel, therefore, deeply interested when we find him making the following declaration: 'The mutual relations of the plants of each great botanical province, and, in fact, of the world generally, is just such as would have resulted if variation had gone on operating throughout indefinite periods, in the same manner as we see it act in a limited number of centuries, so as gradually to give rise in the course of time, to the most widely divergent forms. ...The element of mutability pervades the whole Vegetable Kingdom; no class, nor order, nor genus of more than a few species claims absolute exemption from it, whilst the grand total of unstable forms, generally assumed to be species, probably exceeds that of the stable.'

To many, this doctrine of Natural Selection, or 'the preservation of favoured races in the struggle for life,' seems so simple, when once clearly stated, and so consonant with known facts and received principles, that they have difficulty in conceiving how it can constitute a great step in the progress of science. Such is often the case with important discoveries, but in order to assure ourselves that the doctrine was by no means obvious, we have only to refer back to the writings of skilful naturalists who attempted in the earlier part of the nineteenth century, to theorise on this subject, before the invention of this new method of explaining how certain forms are supplanted by new ones, and in what manner these last are selected out of innumerable varieties, and rendered permanent.

We may understand why the species of the same genus, or genera of the same family, resemble each other more nearly in their embryonic than in their more fully developed state, or how it is that in the eyes of most naturalists the structure of the embryo is even more important in classification than that of the adult, 'for the embryo is the animal in its less modified state, and in so far it reveals the structure of its progenitor. In two groups of animals, however much they may at present differ from each other in structure and habits, if they pass through the same or similar embryonic stages, we may feel assured that they have both descended from the same or nearly similar parents, and are therefore in that degree closely related. Thus community in embryonic structure reveals community of descent, however much the structure of the adult may have been modified.

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The theory of the origin of new species by variation will also explain why a species which has once died out never reappears, and why the fossil fauna and flora recede farther and farther from the living type in proportion as we trace it back to remoter ages. It would also account for the fact, that when we have to intercalate a new set of fossiliferous strata between two groups previously known, the newly discovered fossils serve to fill up gaps between specific or generic types previously familiar to us, supplying often the missing links of the chain, which, if transmutation is accepted, must once have been continuous.

Variation and natural selection would also afford a key to a multitude of geological facts otherwise wholly unaccounted for, as, for example, why there is generally an intimate connection between the living animals and plants of each great division of the globe and the extinct fauna and flora of the post-tertiary or tertiary formations of the same region...

It [variation and natural selection] would explain, says Mr. Darwin, the unity of type which runs through the whole organic world, and why there is sometimes a fundamental agreement in structure in the same class of beings which is quite independent of their habits of life, for such structure, derived by inheritance from a remote progenitor, has been modified, in the course of ages, in different ways, according to the conditions of existence. It would also explain why all living and extinct beings are united, by complex radiating and circuitous lines of affinity with one another, into one grand system; also, there having been a continued extinction of old races and species in progress, and a formation of new ones by variation, why in some genera which are largely represented, or to which a great many species belong, many of these are closely but unequally related; also, why there are distinct geographical provinces of species of animals and plants, for, after long isolation by physical barriers, each fauna and flora, by varying continually, must become distinct from its ancestral type, and from the new forms assumed by other descendants which have diverged from the same stock.

Mr. Darwin labours to show, and with no small success, that all true classification in zoology and botany is, in fact, genealogical, and that community of descent is the hidden bond which naturalists have been unconsciously seeking, while they often imagined that they were looking for some unknown plan of creation.

One of the principal claims of Mr. Darwin's theory to acceptance is, that it enables us to dispense with a law of progression as a necessary accompaniment of variation. It will account equally well for what is called degradation, or a retrograde movement towards a simpler structure, and does not require Lamarck's continual creation of monads; for this was a necessary part of his system, in order to explain how, after the progressive power had been at work for myriads of ages, there were as many beings of the simplest structure in existence as ever.

The competition of races and species, observes Mr. Darwin, is always most severe between those which are most closely allied and which fill nearly the same place in the economy of nature. Hence, when the conditions of existence are modified, the original stock runs great risk of being superseded by some one of its modified offshoots. The new race or species may not be absolutely superior in the sum of its powers and endowment to the parent stock, and may even be more simple in structure and of a lower grade of intelligence, as well as of organisation, provided, on the whole, it happens to have some slight advantage over its rivals. Progression, therefore, is not a necessary accompaniment of variation and natural selection, though, when a higher organisation happens to be coincident with superior fitness to new conditions, the new species will have greater power and a greater chance of permanently maintaining and extending its ground.

Every naturalist admits that there is a general tendency in animals and plants to vary; but it is usually taken for granted, though we have no means of proving the assumption to be true, that there are certain limits beyond which each species cannot pass under any circumstances, or in any number of generations. Mr. Darwin and Mr. Wallace say that the opposite hypothesis, which assumes that every species is capable of varying indefinitely from its original type, is not a whit more arbitrary, and has this manifest claim to be preferred, that it will account for a multitude of phenomena which the ordinary theory is incapable of explaining.

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By the multiplying of slight modifications in the course of thousands of generations, and by the handing down of the newly-acquired peculiarities by inheritance, a greater and greater divergence from the original standard is supposed to be effected, until what may be called a new species, or, in a greater lapse of time, a new genus, will be the result.