The inconsistency of first explaining matter by atoms and then explaining atoms by matter was only slowly realised, and it is only comparatively rece… - Willem de Sitter

Observations give us two data, viz. the rate of expansion and the average density, and there are three unknowns: the value of λ, the sign of the curvature, and the scale of the figure, i.e. the units of R and of the time. The problem is indeterminate.

Matter is actually distributed very unevenly... conglomerated into stars and galactic systems. The average density is the density that we should get if all... could be evaporated into atoms of hydrogen, or protons, and... distributed evenly over the whole of space. ...three or four protons in every cubic foot. ...a million million times less than that of the most perfect vacuum that we can produce... The universe thus consists mostly of emptiness... consider a universe without any matter at all, an empty universe, as a good approximation. But we may also take as our first approximation a universe containing... three or four protons per cubic foot. The local deviations from the average, caused by the conglomeration of matter into stars and stellar systems, are then disregarded in the grand scale model, and are only taken into account when we come to study details.

A question which has long troubled astronomers and physicists is what becomes of the energy that is continually being poured out into space by the sun and the stars. To this question a complete answer is given by the new theory. It is used up, diluted, or degraded, by the expansion of the universe. ...the light travelling through the expanding universe and, so to say, trying to reach a particular star, or stellar system, which is continually receding with great velocity, is losing energy in trying to catch up with it. It is this degradation of the light, technically known as the redshift of the spectral lines, by which we become aware of the receding velocities of the extra-galactic nebulae. It can be shown that the decrease of the total amount of radiant energy in the universe by this degradation exceeds the increase by the radiation of the stars. It would not be correct, however, to conclude that the expansion is caused by the energy thus lost by the radiation...